| Quarter | Holdings | Value ($000) | Form Type | Date Filed | Filing ID | Name |
|---|---|---|---|---|---|---|
| Q1 2023 | 11 | 2,573,424 | 13F-HR | 5/15/2023 | 000095012323005258 | Flagship Pioneering Inc. |
| Q4 2022 | 12 | 3,132,284 | 13F-HR | 2/14/2023 | 000095012323002460 | Flagship Pioneering Inc. |
| Q3 2022 | 12 | 2,375,805 | 13F-HR | 11/14/2022 | 000095012322012261 | Flagship Pioneering Inc. |
| Q2 2022 | 12 | 2,654,624 | 13F-HR | 8/15/2022 | 000095012322009421 | Flagship Pioneering Inc. |
| Q1 2022 | 13 | 4,258,761 | 13F-HR | 5/16/2022 | 000095012322006309 | Flagship Pioneering Inc. |
| Q4 2021 | 13 | 6,467,997 | 13F-HR | 2/14/2022 | 000095012322002901 | Flagship Pioneering Inc. |
| Q3 2021 | 13 | 9,549,130 | 13F-HR | 11/15/2021 | 000095012321015487 | Flagship Pioneering Inc. |
| Q2 2021 | 12 | 7,829,539 | 13F-HR | 8/16/2021 | 000095012321011113 | Flagship Pioneering Inc. |
| Q1 2021 | 12 | 6,244,355 | 13F-HR | 5/17/2021 | 000095012321007079 | Flagship Pioneering Inc. |
| Q4 2020 | 11 | 5,493,282 | 13F-HR | 2/16/2021 | 000095012321002948 | Flagship Pioneering Inc. |
| Q3 2020 | 8 | 3,619,952 | 13F-HR | 11/16/2020 | 000095012320012128 | Flagship Pioneering Inc. |
| Q2 2020 | 8 | 3,441,586 | 13F-HR | 8/13/2020 | 000095012320008652 | Flagship Pioneering Inc. |
| Q1 2020 | 8 | 2,133,438 | 13F-HR | 5/14/2020 | 000095012320004927 | Flagship Pioneering Inc. |
| Q4 2019 | 9 | 1,788,867 | 13F-HR | 2/13/2020 | 000095012320001884 | Flagship Pioneering Inc. |
| Q3 2019 | 10 | 1,828,054 | 13F-HR | 11/14/2019 | 000095012319011393 | Flagship Pioneering Inc. |
| Q2 2019 | 10 | 2,310,279 | 13F-HR | 8/13/2019 | 000095012319007818 | Flagship Pioneering Inc. |
| Q1 2019 | 8 | 2,642,256 | 13F-HR | 5/14/2019 | 000101297519000346 | Flagship Pioneering Inc. |
| Q4 2018 | 7 | 2,047,628 | 13F-HR | 2/13/2019 | 000101297519000145 | Flagship Pioneering Inc. |
| Q3 2018 | 6 | 1,513,015 | 13F-HR | 11/13/2018 | 000101297518001201 | Flagship Pioneering Inc. |
| Q2 2018 | 5 | 522,113 | 13F-HR | 8/7/2018 | 000101297518000952 | Flagship Pioneering Inc. |
| Q1 2018 | 4 | 344,896 | 13F-HR | 5/15/2018 | 000101297518000415 | Flagship Pioneering Inc. |
| Q4 2017 | 5 | 394,322 | 13F-HR | 2/14/2018 | 000101297518000160 | Flagship Pioneering Inc. |
| Q3 2017 | 4 | 298,998 | 13F-HR | 11/3/2017 | 000101297517000806 | Flagship Pioneering Inc. |
| Q2 2017 | 4 | 283,568 | 13F-HR | 8/2/2017 | 000101297517000578 | Flagship Pioneering Inc. |
| Q1 2017 | 7 | 376,254 | 13F-HR | 5/10/2017 | 000101297517000377 | Flagship Pioneering Inc. |
| Q4 2016 | 7 | 307,344 | 13F-HR | 2/13/2017 | 000101297517000113 | Flagship Pioneering Inc. |
| Q3 2016 | 7 | 365,999 | 13F-HR | 11/14/2016 | 000101297516001561 | Flagship Ventures Management, Inc. |